Publications

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Akhmetova, L.A. 2009 A review of scarabs of the subgenus Plagiogonus Mulsant, genus Aphodius Illiger (Coleoptera, Scarabaeidae), of the fauna of Russia and neighboring countries. Entomological Review, 89(4): 428-436.
Akhmetova, L.A., Frolov, A.V. 2014 A review of the scarab beetle tribe Aphodiini (Coleoptera, Scarabaeidae) of the fauna of Russia. Entomological Review, 94, 846-879.
Akhmetova, L.A., Frolov, A.V. 2008 A Review of Scarabs of the Subgenus Nobius Mulsant et Rey, Genus Aphodius Illiger (Coleoptera, Scarabaeidae), of the Fauna of Russia and Neighboring Countries. Entomological Review, 88(4): 421-434.
Akhmetova, L.A., Frolov, A.V. 2012 A new scarab species, Aphodius gissaricus (Coleoptera: Scarabaeidae: Aphodiinae), from the Pamir-Alay Mountains in Tajikistan. Zootaxa, 3159, 65-68.
Akhmetova, L.A., Frolov, A.V. 2009 New to Russia and little known species of the genus Aphodius Illiger (Coleoptera, Scarabaeidae). Zoosystematica Rossica, 18(2): 278–284.
Akhmetova, L.A., Montreuil, O. 2010 Revision of Metadorodocia Machatschke, 1957, a genus endemic to Madagascar (Coleoptera: Scarabaeidae: Rutelinae: Adoretini). Zootaxa, 2401: 61-68.
Davis, A.L.V., Frolov, A.V., Scholtz, C.H. 2008 The African Dung Beetle Genera. Protea Book House, Pretoria. 274 p.
Frolov A. V., Akhmetova L. A., Vaz-de-Mello F. Z. 2017 Revision of the mainland species of the Neotropical genus Aegidium Westwood (Coleoptera: Scarabaeidae: Orphninae). Journal of Natural History. 51 (19–20): 1035–1090.
Frolov A. V., Akhmetova L. A., Vaz-de-Mello F. Z. 2017 Revision of the South American genus Paraegidium (Coleoptera: Scarabaeidae: Orphninae). Journal of Natural History, 51, 995–1014.
Frolov A. V., Akhmetova L. A., Vaz-de-Mello F. Z. 2017 Revision of the Neotropical scarab beetle genus Aegidiellus Paulian (Coleoptera: Scarabaeidae: Orphninae) with description of two new species. Journal of Natural History. 51: 1767–1779.
Frolov A. V., Akhmetova L. A., Vaz-de-Mello F. Z. 2019 Contribution to the knowledge of the Neotropical canopy dwelling genus Onorius (Coleoptera, Scarabaeidae, Orphninae): another species from southern Ecuador and notes on the hind gut morphology in the Orphninae. Journal of Natural History. 53 (17–18): 1071–1077.
Frolov A. V., Akhmetova L. A., Vaz-de-Mello F. Z. 2019 Contribution to the knowledge of Aegidinus Arrow (Coleoptera: Scarabaeidae: Orphninae): new species and comments on the classification and nomenclature. Journal of Natural History. 53 (11-12): 725-747.
Frolov A. V., Montreuil O., Akhmetova L. A. 2016 Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with revision of the genus Triodontus Westwood. Auckland: Magnolia Press. 2016, 93 p.
Frolov A.V. 2013 Little known Orphninae (Coleoptera, Scarabaeidae) collected by the Madagascan expeditions of the California Academy of Sciences. Biodiversity Data Journal, 1, e1016.
Frolov, A. V. & Akhmetova, L. A. 2015 New data on the rare Afrotropical scarab beetles Orphnus drumonti Frolov and Delopleurus naviauxi Frolov et Cambefort (Coleoptera, Scarabaeidae, Orphninae and Scarabaeinae). Biodiversity Data Journal, 3: e5444.
Frolov, A. V. & Akhmetova, L. A. 2015 Rediscovery of the enigmatic Stenosternus costatus Karsch (Coleoptera: Scarabaeidae: Orphninae) from São Tomé island. Zootaxa, 4007, 440–444.
Frolov, A. V. & Akhmetova, L. A. 2015 A new brachypterous scarab species, Orphnus longicornis (Coleoptera: Scarabaeidae: Orphninae), from the East African Rift. Zootaxa, 4039(3): 475–477
Frolov, A. V. & Akhmetova, L. A. 2016 Revision of the subgenus Orphnus (Phornus) (Coleoptera, Scarabaeidae, Orphninae). European Journal of Taxonomy 241: 1–20
Frolov, A. V. & Akhmetova, L. A. 2013 A contribution to the study of the Lower Volga center of scarab beetle diversity in Russia: checklist of the tribe Aphodiini (Coleoptera, Scarabaeidae) of Dosang environs. Biodiversity Data Journal, 1: e979.
Frolov, A.V. 2009 A new species, Orphnus drumonti sp. n. (Coleoptera, Scarabaeidae), from the Democratic Republic of Congo. Entomological Review, 89(6): 753–754.

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